More than 250 fossilized footprints in the Early Jurassic
* Moyeni tracksite located in the Elliot Formation of the Stormberg Group in the Karoo Basin in Lesotho, southern Africa, have been re-examined by researchers using a mix of traditional mapping techniques and a three-dimensional surface scanner, which recorded millimeter-scale detail.
The new analysis of the fossil trackway revealed that two types of dinosaurs, consisting of several basal
ornithischian (bird-hipped) dinosaurs and a single theropod, were walking across an ancient
point bar that presented the animals with a slippery sloping terrain to traverse.
What makes this interesting is the different approaches these two types of dinosaurs took as they adjusted to variations in the terrain as they moved from a wet riverbed to a sloping bank and finally the flat upper surface of the point bar and thus bestowing us a peek at the dynamic locomotive capacities of these two dinosaur lines at an early point in their evolution.
The basal ornithischian track-makers changed the way they walked as surface conditions changed, altering between quadrupedal and bipedal stances, wide and narrow gaits and both plantigrade and digitigrade foot posture.
In the riverbed they appeared to have crouched down low, adopting a sprawling quadrupedal stance, and dragging their feet, crept forward flat-footed. Once on the slope, the ornithischians narrowed their four-legged stance, but no longer dragging their feet. When they finally reached the flat, stable ground on top, they swapped over to walking on only two legs.
The bipedal theropod track-maker, which left around 25 fossilized footprints, was likely a
Grallator that was approximately 5 to 6 meters (16½’ to 19 ½’) long, neither modified foot posture or adopted a wider, more stable gait but rather adjusted to the changing terrain by griping the muddy ground with its pedal claws.
As the researchers noted, the trackway provides a physical record with how two early dinosaurs reacted and adjusted to the same set of complex ground conditions. And the different walking styles also offer insight into the later evolution of these two dinosaur lines.
Ornithischians, which switched from mainly bipedal to exclusively quadrupedal locomotion three separate times in their evolutionary history, are shown to have been capable of adopting both postures early in their evolutionary history.
OTOH, while theropods remained bipedal, the earth-gripping claws are consistent with the hypothesized function of pedal claws in bird ancestors.
Further Reading:
Dynamic Locomotor Capabilities Revealed by Early Dinosaur Trackmakers from Southern Africa Abstract & Paper
Trackway analysis shows how dinosaurs coped with slippery slopes
* The actual age of the sediment the tracks are in are approximately 199 myo – right around the Triassic-Jurassic boundary